1College of Wildlife and Protected Areas, Northeast Forestry University,26, Hexing Road, Xiangfang District, Heilongjiang Province 150040 Harbin, China.
2Tayna Center for Conservation Biology- Kasugho University of Nature Conservation and Development, Goma, D.R.C
3Department of Natural Resources, Egerton University P.O. Box 536-20115 Egerton, Kenya.
4Guangdong Provincial Key Laboratory of Silviculture, Protection and Utilization, Guangdong Academy of Forestry, Guangzhou 510520, China.